3373056 RNLs RPW8-NLRs 2 3373054 RNLs repeat domain CCR -NB-LRR proteins 1 3373055 RNLs RPW8-type HeLo-domain-containing NLRs 1 3373058 RNLs RPW8-type helper NLRs 1 3373057 RNLs Rumex nervosus leaves 1